Publications about Saola

Dawson, S. 1995. Saola Pseudoryx nghetinhensis Studies in Nghe An and Ha Tinh Provinces, Vietnam.  Unpublished report. World Wildlife Fund for  Nature – Indochina Programme, National Wildlife Federation and World Conservation Union, Hanoi.

Dung, V. V.,  Giao, P.M.,  Chinh, N. N., Touc D., Arctander P., and MacKinnon, J.  1993.  “A new species of living bovid from Vietnam.Nature, 363:443-445.

Dung, V. V., Giao, P. M., Chinh, N. N., Tuoc, D. and MacKinnon J. 1994. “Discovery and conservation of the Vu Quang ox in Vietnam.”  Oryx28:16-21.

Dung, V. V., Nguyen, N. C., Ebregt, A. and Santiapillai, C. 1995.  “The status of the newly discovered large mammal, the long-horned bovid, in Vietnam.”  TigerPaper, 22:13-19.

Gatesy, J. and Arctander, P. 2000. “Hidden morphological support for the phylogenetic placement of Pseudoryx nghetinhensis with bovine bovids: a combined analysis of gross anatomical evidence and DNA sequences from five genes.” Systematic Biology 49: 515-538.

Hardcastle, J., Cox, S., Dao, N. T. and Johns, A. G. 2004. Rediscovering the Saola – Workshop Proceedings. WWF Indochina Programme, SFNC Project, Pu Mat National Park.

Hassanin, A. and Douzery, E. J. P. 1999. “Evolutionary affinities of the enigmatic Saola (Pseudoryx nghetinhensis) in the context of the molecular phylogeny of Bovidae.Proc. R. Soc. Lond. B Biol. Sci. 266: 893-900.

Henshaw, J. 1997.  “Status of Saola Pseudoryx nghetinhensis.” Oryx, 31: 89-91.

Huynh, D.H., Anh, P.T., Khien, H.M., Canh, L.X., Thang, T.V. and Phuong, D.H. 1997. “Supplemental data on distribution of Saola Pseudoryx nghetinhensis in Vietnam.” Tap Chi SINH HOC 19:16-19 (in Vietnamese; English translation by FFI Indochina Programme).

Kemp, N., Dilger, M., Burgess N., and Dung, C. V. 1997.  “The Saola Pseudoryx nghetinhensis in Vietnam – new information on distribution and habitat preferences, and conservation needs.” Oryx, 31: 37-44.

Khanh, H.N., Tin, N.V.T. and Tuan, N.D.A. 1998. “Results of a Survey for Saola (Pseudoryx nghetinhensis) in Thua Thien Hue Province.”  Thua Thien-Hue Province Forestry Management Survey Section, Hue, Vietnam (in Vietnamese)

Nguyen, T. T., et al. “Application of bovine microsatellite markers on Saola (Pseudoryx nghetinhensis).” Journal of Animal Breeding and Genetics 122.3 (2005): 195-198.

Nguyen, T. T., et al. “Phylogenetic position of the saola (Pseudoryx nghetinhensis) inferred from cytogenetic analysis of eleven species of Bovidae.” Cytogenetic and genome research 122.1 (2008): 41-54.

Oanh, D. C. 2004. “Real situation and solutions of conservation of Saola in Pumat National Park.” Nong Nghiep Va Phat Trien Nong Thon (in Vietnamese).

Phaengsintham, P. 1996.  Survey of Food Plants and Saltlicks Used by Saola (Pseudoryx nghetinhensis). Dong Dok Pedagogical University and Wildlife Conservation Society, Vientiane.

Proschan, F. 1994. The /saavlaaa/ (Pseudoryx nghetinhensis). Research Center for Language and Semiotic Studies, Indiana University, Bloomington, USA.

Provincial Forest Protection Department of Quang Nam-Da Nang. 1996. Results of a Preliminary Survey on the Saola, Pseudoryx nghetinhensis, in Quang Nam-Da Nang Province. Provincial Forest Protection Department of Quang Nam-Da Nang (in Vietnamese with English translation).

Reinecke, A. 1996. Bi-Cephalous Animal-Shaped Ear Pendants in Vietnam. BSTN, 28: 5-8

Robichaud, W.G. 1998a. Physical and behavioral description of a captive Saola Pseudoryx nghetinhensis.  J. Mamm., 79: 394-405.

Robichaud, W. 1998b.  WCS trip report:  Bolikhamxay Saola Survey, 15-24 January 1998. Internal trip report. Wildlife Conservation Society, Vientiane.

Robichaud, W. 1999. Saola Conservation Action Plan for Lao PDR: 1999 Revision. Wildlife Conservation Society and IUCN. Vientiane.

Robichaud, W. and Stuart, B.L. 1999. Saola, Herpetological and Wildlife Trade Studies in Nakai-Nam Theun National Biodiversity Conservation Area and the Proposed Nam Theun Extension. Wildlife Conservation Society and IUCN. Vientiane.

Robichaud, W. and Timmins, R. 2004. The Natural History of Saola (Pseudoryx nghetinhensis) and the Species’ Distribution in Laos. Presentation given at the symposium “Rediscovering the Saola: A status review and conservation planning workshop”, held at Pu Mat Nature Reserve, Vietnam, 27-28 February 2004.

Schaller, G. B. and Rabinowitz, A.  1995.  “The Saola or spindlehorn bovid Pseudoryx nghetinhensis in Laos.” Oryx, 29:107-113.

Souvannalath, P. 1998. [Wildlife Survey (Saola, or “Nyu-ja”) From Villages in Bolikhan District, Bolikhamxay Province]. Unpublished report to WCS.  Bolikhamxay Province Forestry Office, Pakxan, Lao PDR (in Lao).

Stone, Richard. “Last Chance to Save the ‘Panda of Indochina’.” Science 325.5945 (2009): 1192-1193.

Thomas,  H.  1994.  “Anatomie crânniene et relations phylogénétiques du nouveau bovidé Pseudoryx nghetinhensis découvert dans la cordillère annamatique au Vietnam.”  Mammalia, 58:453-481.

Tilker, A. 2014.  “A survey of eastern areas of Xe Sap National Protected Area, Lao PDR, for Saola and other large ungulates; final report to Global Wildlife Conservation and the Saola Working Group.” World Wildlife Fund Greater Mekong Program. Vientiane.

Timmins, R. J. 2001.  Chapter 37: Lao PDR,  pp 194-199 in: Mallon, D. P. and Kingswood, S. C. (compilers). Antelopes. Part 4: North Africa, the Middle East, and Asia. Global survey and regional Action Plans. SSC Antelope Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK.

Tordoff, Andrew W. 2003. A biological assessment of the Central Truong Son Landscape. (No. 1). WWF Indochina, Hanoi.

Turvey, S. T., Trung, C. T., Quyet, V. D., Nhu, H. V., Thoai, D. V., Tuan, V. C. A., Hoa, D. T., Kacha, K., Sysomphone, T., Wallate, S., Hai, C. T. T., Thanh, N. V., Wilkinson, N. M. 2015. “Interview-based sighting histories can inform regional conservation prioritization for highly threatened cryptic species.” Journal of Applied Ecology, 52: 422–433.

Please contribute to this list. Mail further references, ideally with the corresponding PDF, to